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I am relatively new to NLME, so the solution to the problem I describe here may be obvious. But I've spent several days trying to get the right syntax to formulate random effects for this model appropriately. The full model is: nlme(a ~ a.mitscherlich(a.qe, a.max, lcp, light), data=light, fixed = a.max + a.qe + lcp ~ trt, random = a.max + a.qe + lcp ~ 1 | bench/line,